Wang, Junfeng, School of Environmental Science and Technology, Shanxi University of Science & Technology, Xi’an, Shanxi, 710021,China;State Key Laboratory of Frozen Soil Engineering, Cold and Arid Regions Environmental and Engineering Research Institute, Chinese Academy of Sciences, Lanzhou, Gansu, 730000, China;Department of Microbiology and Plant Biology, University of Oklahoma, Norman, OK 73019, USA;Beiluhe Observation Station of Frozen Soil Environment and Engineering, Cold and Arid Regions Environmental and Engineering Research Institute, Chinese Academy of Sciences, Lanzhou, Gansu, 730000, China