Assou, Délagnon, Laboratory of Hydrobiology and Aquaculture (LHA), Faculty of Agricultural Sciences, University of Abomey-Calavi, 01BP 526 Cotonou-Benin;Laboratory of Ecology and Ecotoxicology (LaEE), Department of Zoology and Animal Biology, Faculty of Sciences, University of Lomé, 01BP 1515 Lomé-Togo;Togolese Society for Nature Conservation (AGBO-ZEGUE NGO), 06 BP: 6057 Lomé-Togo