Zhou, Guowei, Key Laboratory of Tropical Marine Bio-resources and Ecology, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, 510301, China; Innovation Academy of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences,Guangzhou, 510301, China; Tropical Marine Biological Research Station in Hainan, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Sanya, 572000, China