Li, Jiezhang, College of Animal Science, Guizhou University, Guiyang 550025, Guizhou, China;Key Laboratory of Genetics, Breeding and Reproduction of Plateau and Mountain Animals of the Ministry of Education, Guizhou University, Guiyang 550025, Guizhou, China;Guizhou Province Key Laboratory of Animal Genetics, Breeding and Reproduction, Guizhou University, Guiyang 550025, Guizhou, China